LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

The Galápagos Islands are famous for their biodiversity, especially their giant tortoises. But around 180 years ago Floreana Island lost its unique tortoise, the Floreana giant tortoise (Chelonoidis ...